Konrad Lorenz

Konrad Zacharias Lorenz (Austrian German: [ˈkɔnraːd tsaxaˈriːas ˈloːrɛnts] ; 7 November 1903 – 27 February 1989) was an Austrian zoologist, ethologist, and ornithologist. He shared the 1973 Nobel Prize in Physiology or Medicine with Nikolaas Tinbergen and Karl von Frisch. He is often regarded as one of the founders of modern ethology, the study of animal behavior. He developed an approach that began with an earlier generation, including his teacher Oskar Heinroth.

Lorenz studied instinctive behavior in animals, especially in greylag geese and jackdaws. Working with geese, he investigated the principle of imprinting, the process by which some nidifugous birds (i.e. birds that leave their nest early) bond instinctively with the first moving object that they see within the first hours of hatching. Although Lorenz did not discover the topic, he became widely known for his descriptions of imprinting as an instinctive bond. In 1936, he met Tinbergen, and the two collaborated in developing ethology as a separate sub-discipline of biology. A Review of General Psychology survey, published in 2002, ranked Lorenz the 65th most cited scholar of the 20th century in the technical psychology journals, introductory psychology textbooks, and survey responses.

Lorenz's work was interrupted by the onset of World War II, and in 1941 he was recruited into the German Army as a medic. In 1944, he was sent to the Eastern Front, where he was captured by the Soviet Red Army and spent four years as a German prisoner of war in Soviet Armenia. After the war, he regretted his membership in the Nazi Party.

Lorenz wrote numerous books, some of which, such as King Solomon's Ring, On Aggression, and Man Meets Dog, became popular reading. His last work Here I Am – Where Are You? is a summary of his life's work and focuses on his famous studies of greylag geese.

Lorenz was the son of Adolf Lorenz, a wealthy and distinguished surgeon, and his wife Emma (née Lecher), a physician who had been her husband's assistant. The family lived on a large estate at Altenberg and had a city apartment in Vienna. He was educated at the Public Schottengymnasium of the Benedictine monks in Vienna.

In his autobiographical essay, published in 1973 in Les Prix Nobel (winners of the prizes are requested to provide such essays), Lorenz credits his career to his parents, who "were supremely tolerant of my inordinate love for animals", and to his childhood encounter with Selma Lagerlöf's The Wonderful Adventures of Nils, which filled him with a great enthusiasm about wild geese.

At the request of his father, he began a premedical curriculum in 1922 at Columbia University, but returned to Vienna in 1923 to continue his studies at the University of Vienna. He graduated as Doctor of Medicine (MD) in 1928 and became an assistant professor at the Institute of Anatomy until 1935. He finished his zoological studies in 1933 and received his second doctorate (PhD).

While still a student, Lorenz began developing what would become a large menagerie, ranging from domestic to exotic animals. In his popular book King Solomon's Ring, Lorenz recounts that while studying at the University of Vienna, he kept a variety of animals at his parents' apartment, ranging from fish to a capuchin monkey named Gloria.

In 1936, at an international scientific symposium on instinct, Lorenz met his great friend and colleague Nikolaas Tinbergen. Together they studied geese – wild, domestic, and hybrid. One result of these studies was that Lorenz "realized that an overpowering increase in the drives of feeding as well as of copulation and a waning of more differentiated social instincts is characteristic of very many domestic animals". Lorenz began to suspect and fear "that analogous processes of deterioration may be at work with civilized humanity". This observation of bird hybrids caused Lorenz to believe that domestication resulting from urbanisation in humans might also cause dysgenic effects, and to argue in two papers that the Nazi eugenics policies against this were therefore scientifically justified.

Lorenz wed his childhood companion Margarethe Gebhardt, a gynaecologist and the daughter of a local market gardener. The couple had three children – one son and two daughters. He resided on the Lorenz family estate, notable for its "fantastical neo-baroque mansion", formally belonging to his father.

Lorenz is recognized as one of the founding fathers of the field of ethology, the study of animal behavior. He is best known for his research of the principle of attachment, or imprinting, through which in some species a bond is formed between a newborn animal and its caregiver. This principle had been discovered by Douglas Spalding in the 19th century, and Lorenz's mentor Oskar Heinroth had also worked on the topic, but Lorenz's description of Prägung, imprinting, in nidifugous birds such as greylag geese in his 1935 book Der Kumpan in der Umwelt des Vogels ("The Companion in the Environment of Birds") became the foundational description of the phenomenon.

Here, Lorenz used Jakob von Uexküll's concept of Umwelt to understand how the limited perception of animals filtered out certain phenomena with which they interacted instinctively. For example, a young goose instinctively bonds with the first moving stimulus it perceives, whether it be its mother, or a person. Lorenz showed that this behavior of imprinting is what allows the goose to learn to recognize members of its own species, enabling them to be the object of subsequent behavior patterns such as mating. He developed a theory of instinctive behavior that saw behavior patterns as largely innate but triggered through environmental stimuli, for example, the hawk/goose effect. He argued that animals have an inner drive to carry out instinctive behaviors, and that if they do not encounter the right stimulus, they will eventually engage in the behavior with an inappropriate stimulus.

Lorenz's approach to ethology derived from a skepticism towards the studies of animal behavior done in laboratory settings. He considered that in order to understand the mechanisms of animal behavior, it was necessary to observe their full range of behaviors in their natural context. Lorenz did not carry out much traditional fieldwork but observed animals near his home. His method involved empathizing with animals, often using anthropomorphization to imagine their mental states. He believed that animals were capable of experiencing many of the same emotions as humans.

Tinbergen, Lorenz's friend with whom he conjointly received the Nobel Prize, summarized Lorenz's major contribution to ethology as making behavior a topic of biological inquiry, considering behavior a part of an animal's evolutionary equipment. Tinbergen and Lorenz contributed to making ethology a recognized sub-discipline within biology and founded the first specialized journal of the field "Ethology" (originally Zeitschift für Tierpsychologie).

Lorenz joined the Nazi Party in 1938 and accepted a university chair under the Nazi regime. In his application for party membership he wrote: "I'm able to say that my whole scientific work is devoted to the ideas of the National Socialists." His publications during that time led in later years to allegations that his scientific work had been contaminated by Nazi sympathies. His published writing during the Nazi period included support for Nazi ideas of "racial hygiene" couched in pseudoscientific metaphors.

The same individual geese on which we conducted these experiments, first aroused my interest in the process of domestication. They were F1 hybrids of wild Greylags and domestic geese and they showed surprising deviations from the normal social and sexual behaviour of the wild birds. I realised that an overpowering increase in the drives of feeding as well as of copulation and a waning of more differentiated social instincts is characteristic of very many domestic animals. I was frightened – as I still am – by the thought that analogous genetical processes of deterioration may be at work with civilized humanity. Moved by this fear, I did a very ill-advised thing soon after the Germans had invaded Austria: I wrote about the dangers of domestication and, in order to be understood, I couched my writing in the worst of Nazi-terminology. I do not want to extenuate this action. I did, indeed, believe that some good might come of the new rulers. The precedent narrow-minded Catholic regime in Austria induced better and more intelligent men than I was to cherish this naive hope. Practically all my friends and teachers did so, including my own father who certainly was a kindly and humane man. None of us as much as suspected that the word "selection", when used by these rulers, meant murder. I regret those writings not so much for the undeniable discredit they reflect on my person as for their effect of hampering the future recognition of the dangers of domestication.

After the war, Lorenz denied having been a party member, until his membership application was made public, and he denied having known the extent of the genocide, despite his position as a psychologist in the Office of Racial Policy. He was shown to have made anti-Semitic jokes on "Jewish characteristics" in letters to his mentor Heinroth. In 2015, the University of Salzburg posthumously rescinded an honorary doctorate awarded to Lorenz in 1983, citing his party membership and his assertions in his application that he was "always a National Socialist", and that his work "stands to serve National Socialist thought". The university also accused him of using his work to spread "basic elements of the racist ideology of National Socialism".

During the final years of his life, Lorenz supported the fledgling Austrian Green Party and in 1984 became the figurehead of the Konrad Lorenz Volksbegehren, a grass-roots movement that was formed to prevent the building of a power plant at the Danube near Hainburg an der Donau and thus the destruction of the surrounding woodland.

Lorenz has been called "The father of ethology" by Niko Tinbergen. Perhaps Lorenz's most important contribution to ethology was his idea that behavior patterns can be studied as anatomical organs. This concept forms the foundation of ethological research. However, Richard Dawkins called Lorenz a "'good of the species' man", stating that the idea of group selection was "so deeply ingrained" in Lorenz's thinking that he "evidently did not realize that his statements contravened orthodox Darwinian theory".

Together with Nikolaas Tinbergen, Lorenz developed the idea of an innate releasing mechanism to explain instinctive behaviors (fixed action patterns). They experimented with "supernormal stimuli" such as giant eggs or dummy bird beaks, which they found could release the fixed action patterns more powerfully than the natural objects for which the behaviors were adapted. Influenced by the ideas of William McDougall, Lorenz developed this into a "psychohydraulic" model of the motivation of behavior, which tended towards group selectionist ideas, which were influential in the 1960s. Another of his contributions to ethology is his work on imprinting. His influence on a younger generation of ethologists and his popular works were important in bringing ethology to the attention of the general public.

Lorenz claimed that there was widespread contempt for the descriptive sciences. He attributed this to the denial of perception as the source of all scientific knowledge: "a denial that has been elevated to the status of religion". He wrote that in comparative behavioral research, "it is necessary to describe various patterns of movement, record them, and above all, render them unmistakably recognizable".

There are three research institutions named after Lorenz in Austria: the Konrad Lorenz Institute for Evolution and Cognition Research (KLI) was housed in Lorenz' family mansion at Altenberg before moving to Klosterneuburg in 2013; the Konrad Lorenz Forschungsstelle (KLF) at his former field station in Grünau; and the Konrad Lorenz Institute of Ethology, an external research facility of the University of Veterinary Medicine Vienna.

Lorenz predicted the relationship between market economics and the threat of ecological catastrophe. In his 1973 book Civilized Man's Eight Deadly Sins, Lorenz addresses the following paradox:

All the advantages that man has gained from his ever-deepening understanding of the natural world that surrounds him, his technological, chemical and medical progress, all of which should seem to alleviate human suffering ... tends instead to favor humanity's destruction.

Lorenz adopts an ecological model to attempt to grasp the mechanisms behind this contradiction. Thus "all species ... are adapted to their environment ... including not only inorganic components ... but all the other living beings that inhabit the locality".

Fundamental to Lorenz's theory of ecology is the function of negative feedback mechanisms, which, in hierarchical fashion, dampen impulses that occur beneath a certain threshold. The thresholds themselves are the product of the interaction of contrasting mechanisms. Thus pain and pleasure act as checks on each other:

To gain a desired prey, a dog or wolf will do things that, in other contexts, they would shy away from: run through thorn bushes, jump into cold water and expose themselves to risks which would normally frighten them. All these inhibitory mechanisms ... act as a counterweight to the effects of learning mechanisms ... The organism cannot allow itself to pay a price which is not worth paying.

In his 1973 book Behind the Mirror: A Search for a Natural History of Human Knowledge, Lorenz considers the old philosophical question of whether our senses correctly inform us about the world as it is, or provide us only with an illusion. His answer comes from evolutionary biology. Only traits that help us survive and reproduce are transmitted. If our senses gave us wrong information about our environment, we would soon be extinct. Therefore, we can be sure that our senses give us correct information, for otherwise we would not be here to be deceived.

Elected a member of the American Academy of Arts and Sciences in 1957

Austrian Decoration for Science and Art in 1964

Elected a Foreign Member of the Royal Society (ForMemRS) in 1964

Elected a member of the United States National Academy of Sciences in 1966

Kalinga Prize for the Popularization of Science in 1969

Lorenz's best-known books are King Solomon's Ring and On Aggression, both written for a popular audience. His scientific work appeared mainly in journal articles, written in German; it became widely known to English-speaking scientists through its description in Tinbergen's 1951 book The Study of Instinct, though many of his papers were later published in English translation in the two volumes titled Studies in Animal and Human Behavior.

King Solomon's Ring (1949) (Er redete mit dem Vieh, den Vögeln und den Fischen, 1949)

Man Meets Dog (1950) (So kam der Mensch auf den Hund, 1950)

Evolution and Modification of Behaviour (1965)

On Aggression (1966) (Das sogenannte Böse. Zur Naturgeschichte der Aggression, 1963)